Greater roadrunners measure 20 to 24 inches (52 to 62 cm) from beak to tail, with the tail accounting for roughly half that length. They weigh 8 to 15 ounces (227 to 425 grams). Plumage is brown, white, and black streaked — excellent camouflage against desert gravel and brush.
Strong legs and zygodactyl feet (two toes forward, two back) grip the ground for explosive sprints reaching 20 mph (32 km/h). A shaggy blue-black crest can be raised or lowered. The long tail acts as a rudder during sharp turns.
Roadrunners can lower their body temperature at night to conserve energy, then sunbathe in the morning with back feathers spread to absorb heat — a behavior called dorsal sunning.