Hippos are colossally heavy, bulls larger than cows: a mature male runs 10 to 16 feet long, up to 5.5 feet at the shoulder, and 3,300 to over 4,400 pounds, yet can hit 19 mph in a short charge on land. The skull is built for amphibious life, with eyes, ears, and nostrils set high on top so the animal can see, hear, and breathe while almost fully submerged, and muscular valves that seal the ears and nostrils when it dives. The skin is up to two inches thick as armor against rival tusks, but lacks sweat glands, so hippos secrete a viscous red "blood sweat" — neither blood nor sweat, but a glandular fluid that works as sunscreen and antibacterial protection in pathogen-rich water. The jaws open to a huge 150-to-180-degree gape, revealing self-sharpening ivory canines up to 20 inches long. It belongs to the family Hippopotamidae; surprisingly, genomics shows the hippo's closest living relatives are not pigs but the cetaceans — whales and dolphins — from a common semi-aquatic ancestor around 55 million years ago.